Philippine Journal of Natural Sciences

The Philippine Journal of Natural Sciences (formerly UPV Journal of Natural Sciences) publishes original articles, short communications, and review papers on problems and issues in broad fields of Biology, Physics, Chemistry, Mathematics, and in Fisheries Science including Aquaculture.

Volume 26, No 1 & 2 (2021)

EXECUTIVE EDITOR

Mae Grace G. Nillos

Institute of Aquaculture, College of Fisheries and Ocean Sciences,
College of Fisheries and Ocean Sciences,
University of the Philippines Visayas

MANAGING EDITOR

Harold M. Monteclaro, Ph.D.
Institute of Marine Fisheries and Oceanology,
College of Fisheries and Ocean Sciences,
University of the Philippines Visayas

INVITED EDITORIAL BOARD MEMBERS

Philip Ian P. Padilla, Ph.D.
Division of Biological Sciences,
College of Arts and Sciences,
University of the Philippines Visayas, Miagao, Iloilo
Andrea Roxanna Jocsing-Anas, Ph.D.
Research Institute of Environmental Medicine,
Nagoya University,
Japan
Mary Jane A. Amar, Ph.D.
College of Fisheries and Ocean Sciences,
University of the Philippines Visayas,
Miagao, Iloilo
Teddy Yap, Ph.D.
Algonquin College,
Ontario, Canada
Peter Paolo Rivera, Ph.D.
Scottish Association for Marine Sciences,
United Kingdom
Irma Tandingan De Ley, Ph.D.
College of Natural and Agricultural Sciences,
University of California,
Riverside, USA
Jhoanne Marsh Gatpatan, Ph.D.
Division of Physical Sciences and Mathematics,
College of Arts and Sciences,
University of the Philippines Visayas, Miagao, Iloilo
Eric Zeus C. Rizo, Ph.D.
Division of Biological Sciences,
College of Arts and Sciences,
University of the Philippines Visayas, Miagao, Iloilo

Articles

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UP
UPV

University of the Philippines Visayas

Office of the Vice Chancellor for Research and Extensions

Miagao, Iloilo

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ovcre.upvisayas@up.edu.ph

(02) 315-8382